See: Genetic differentiation without mimicry shift in a pair of hybridizing Heliconius species (Lepidoptera: Nymphalidae) CLAIRE MÉROT, JESÚS MAVÁREZ, ALLOWEN EVIN,KANCHON K. DASMAHAPATRA, JAMES MALLET, GERARDO LAMAS and MATHIEU JORON1″ – Biological Journal of the Linnean Society, 2013